"the source of the effect remains unexplained" They list several candidate explanations, but miss one I came up with immediately: maybe zebras are in some other way less attractive targets for the flies -- their skin is thicker, their blood/fluids don't taste as good, etc. -- and the stripes are simply a visual indicator to the flies that they are zebras, and therefore less desirable. This sort of trait has evolved poison frogs, insects, and other plants and animals, so why not zebras?